Structural Plasticity of PAM Recognition by Engineered Variants of the RNA-Guided Endonuclease Cas9

Mar 19, 2016Molecular cell

Changes in DNA Target Recognition by Modified Versions of the Gene-Editing Protein Cas9

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Abstract

Crystal structures of SpCas9 variants reveal an induced fit mechanism for recognizing non-canonical PAMs.

  • DNA cleavage by SpCas9 requires a specific 5'-NGG-3' protospacer adjacent motif (PAM), limiting target sequences.
  • Artificial SpCas9 variants with altered PAM specificities have been developed to overcome these limitations.
  • The structures of VQR, EQR, and VRER variants bound to their preferred PAM sequences show conformational changes in the PAM region of the DNA.
  • Amino acid substitutions in these SpCas9 variants enhance sequence-specific base recognition and conformational remodeling.
  • A new SpCas9 variant was engineered to specifically recognize NAAG PAMs based on structural insights.

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