Rhythmic E-Box Binding by CLK-CYC Controls Daily Cycles in per and tim Transcription and Chromatin Modifications

May 14, 2008Molecular and cellular biology

Daily cycles of per and tim gene activity and DNA packaging controlled by CLK-CYC binding rhythm

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Abstract

Rhythmic transcription of the per and tim genes involves time-of-day-specific binding of CLK-CYC and associated chromatin modifications.

  • Activation of per and tim transcription occurs alongside CLK-CYC binding to E-boxes and specific histone modifications.
  • Histone H3-K9 acetylation and H3-K4 trimethylation are observed during the activation phase of these genes.
  • RNA polymerase II binding to the tim promoter is linked to transcriptional elongation, while it remains constantly bound to the per promoter.
  • Repression of transcription is associated with the reversal of these modifications by PER.
  • Similar rhythmic histone modifications are linked to CLOCK-BMAL1-dependent transcription in mammals, suggesting a conserved mechanism across species.

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