KaiA-stimulated KaiC phosphorylation in circadian timing loops in cyanobacteria

Oct 23, 2002Proceedings of the National Academy of Sciences of the United States of America

KaiA helps activate KaiC to keep the biological clock running in cyanobacteria

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Abstract

Activation of kaiBC expression by kaiA requires KaiC, indicating a positive feedback mechanism in the cyanobacterial circadian clock.

  • KaiA is identified as a positive regulator of kaiBC expression.
  • KaiC acts as a negative regulator in this autoregulatory system.
  • Robust circadian phosphorylation of KaiC was observed.
  • KaiA is essential for both in vivo phosphorylation of KaiC and in vitro autophosphorylation of KaiC.
  • A long period mutation in kaiA2 attenuated the effects of KaiA on KaiC phosphorylation.
  • A previously undocumented mutation in kaiC suppressed the long period phenotype and abnormal phosphorylation in the kaiA2 mutant.

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